About the Authors

Ryusuke Matsuda

Contributed equally to this work with: Ryusuke Matsuda, Julio Cesar Masaru Iehisa

Affiliation Laboratory of Plant Genetics, Graduate School of Agricultural Science, Kobe University, Kobe, Japan

Julio Cesar Masaru Iehisa

Contributed equally to this work with: Ryusuke Matsuda, Julio Cesar Masaru Iehisa

Affiliation Departmento de Biotecnología, Facultad de Ciencias Químicas, Universidad Nacional de Asunción, San Lorenzo, Paraguay

Kouhei Sakaguchi

Affiliation Laboratory of Plant Genetics, Graduate School of Agricultural Science, Kobe University, Kobe, Japan

Ryoko Ohno

Affiliation Graduate School of Science, Technology and Innovation, Kobe University, Kobe, Japan

Kentaro Yoshida

Affiliation Laboratory of Plant Genetics, Graduate School of Agricultural Science, Kobe University, Kobe, Japan

Shigeo Takumi

takumi@kobe-u.ac.jp

Affiliation Laboratory of Plant Genetics, Graduate School of Agricultural Science, Kobe University, Kobe, Japan

Competing Interests

The authors partly received funding from a commercial source of KANEKA Co. Ltd. However, there is no competing interest relating to employment, consultancy, patents, products in development, marketed products, etc. in this article. This does not alter our adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceptualization: ST. Data curation: RM JCMI KY. Formal analysis: JCMI KY. Funding acquisition: ST. Investigation: RM JCMI KS RO. Project administration: ST. Resources: ST. Supervision: ST. Validation: JCMI KY ST. Visualization: JCMI KY ST. Writing – original draft: ST. Writing – review & editing: JCMI KY ST.