About the Authors

Philippa K. Morrison

Affiliation University of Surrey, School of Veterinary Medicine, Faculty of Health and Medical Sciences, Guilford, United Kingdom

Patricia A. Harris

Affiliation Equine Studies Group, WALTHAM Centre for Pet Nutrition, Freeby Lane, Waltham-on-the-Wolds, Melton Mowbray, Leicestershire, United Kingdom

Charlotte A. Maltin

Affiliations University of Liverpool, Department of Obesity and Endocrinology, Faculty of Health and Life Sciences, Leahurst Campus, Chester High Road, Neston, Wirral, United Kingdom, Biomics Ltd, Inverurie, Aberdeenshire, United Kingdom

Dai Grove-White

Affiliation University of Liverpool, Department of Obesity and Endocrinology, Faculty of Health and Life Sciences, Leahurst Campus, Chester High Road, Neston, Wirral, United Kingdom

Caroline McG. Argo

c.argo@surrey.ac.uk

Affiliation University of Surrey, School of Veterinary Medicine, Faculty of Health and Medical Sciences, Guilford, United Kingdom

Competing Interests

Patricia A. Harris is employed by one of the funders of this research (WALTHAM Centre for Pet Nutrition, Melton Mowbray, Leicestershire, United Kingdom). The authors can confirm that they have adhered to all the PLOS ONE policies on sharing data and materials. The authors also confirm that none of the authors record a conflict of interest.

Author Contributions

Conceptualization: PM PH CM DGW CMA. Data curation: PM CMA. Formal analysis: PM DGW CMA. Investigation: PM CMA. Methodology: PM PH CM DGW CMA. Writing – original draft: PM CMA CM DGW. Writing – review & editing: PM PH CM DGW CMA.