About the Authors

Pieter I. Olivier

polivier@zoology.up.ac.za

Affiliation Conservation Ecology Research Unit, Department of Zoology and Entomology, University of Pretoria, Pretoria, Hatfield, South Africa

Victor Rolo

Affiliation Conservation Ecology Research Unit, Department of Zoology and Entomology, University of Pretoria, Pretoria, Hatfield, South Africa

Rudi J. van Aarde

Affiliation Conservation Ecology Research Unit, Department of Zoology and Entomology, University of Pretoria, Pretoria, Hatfield, South Africa

Competing Interests

We have been funded in part by a commercial source, Richards Bay Minerals. This does not alter our adherence to all PLOS ONE policies on sharing data and materials.

Author Contributions

Conceptualization: PIO RJV. Data curation: PIO. Formal analysis: VR PIO. Funding acquisition: RJV PIO VR. Investigation: PIO. Methodology: VR PIO. Project administration: PIO RJV. Software: VR. Supervision: RJV. Visualization: PIO VR RJV. Writing — original draft: PIO. Writing — review & editing: PIO VR RJV.