About the Authors

Lin Li

Contributed equally to this work with: Lin Li, Jinko Sawashita

Affiliation Department of Aging Biology, Institute of Pathogenesis and Disease Prevention, Shinshu University Graduate School of Medicine, Matsumoto, Japan

Jinko Sawashita

Contributed equally to this work with: Lin Li, Jinko Sawashita

jinkos@shinshu-u.ac.jp

Affiliations Department of Aging Biology, Institute of Pathogenesis and Disease Prevention, Shinshu University Graduate School of Medicine, Matsumoto, Japan, Department of Biological Sciences for Intractable Neurological Diseases, Institute for Biomedical Sciences, Interdisciplinary Cluster for Cutting Edge Research, Shinshu University, Matsumoto, Japan

Xin Ding

Affiliation Department of Aging Biology, Institute of Pathogenesis and Disease Prevention, Shinshu University Graduate School of Medicine, Matsumoto, Japan

Mu Yang

Affiliation Department of Aging Biology, Institute of Pathogenesis and Disease Prevention, Shinshu University Graduate School of Medicine, Matsumoto, Japan

Zhe Xu

Affiliation Department of Aging Biology, Institute of Pathogenesis and Disease Prevention, Shinshu University Graduate School of Medicine, Matsumoto, Japan

Hiroki Miyahara

Affiliation Department of Aging Biology, Institute of Pathogenesis and Disease Prevention, Shinshu University Graduate School of Medicine, Matsumoto, Japan

Masayuki Mori

Affiliations Department of Aging Biology, Institute of Pathogenesis and Disease Prevention, Shinshu University Graduate School of Medicine, Matsumoto, Japan, Department of Advanced Medicine for Health Promotion, Institute for Biomedical Sciences, Interdisciplinary Cluster for Cutting Edge Research, Shinshu University, Matsumoto, Japan

Keiichi Higuchi

Affiliations Department of Aging Biology, Institute of Pathogenesis and Disease Prevention, Shinshu University Graduate School of Medicine, Matsumoto, Japan, Department of Biological Sciences for Intractable Neurological Diseases, Institute for Biomedical Sciences, Interdisciplinary Cluster for Cutting Edge Research, Shinshu University, Matsumoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: JS KH. Formal analysis: JS HM. Funding acquisition: JS KH. Investigation: LL JS XD MY ZX HM. Methodology: LL JS KH. Project administration: JS KH. Resources: LL JS MM KH. Software: HM. Supervision: JS KH. Validation: JS. Visualization: LL JS. Writing – original draft: LL JS KH. Writing – review & editing: JS KH.