About the Authors

Toshiyuki Komatsu

Affiliation Graduate School of Agriculture, Kyoto University, Kyoto, Japan

Suguru Sasaki

Affiliation Graduate School of Agriculture, Kyoto University, Kyoto, Japan

Yuki Manabe

Affiliation Graduate School of Agriculture, Kyoto University, Kyoto, Japan

Takashi Hirata

Current address: Shijonawate Gakuen University, Daito, Osaka, Japan

Affiliation Graduate School of Agriculture, Kyoto University, Kyoto, Japan

Tatsuya Sugawara

sugawara@kais.kyoto-u.ac.jp

Affiliation Graduate School of Agriculture, Kyoto University, Kyoto, Japan

Competing Interests

FUJIFILM Corporation funded this study and provided support for the author TS. The other authors state no conflict of interest. This does not alter our adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceptualization: TS TH. Formal analysis: TK SS TS. Funding acquisition: TS. Investigation: TK SS YM. Project administration: TH TS. Writing – original draft: TS. Writing – review & editing: TK SS YM TH TS.