About the Authors

Sharavari S. Kulkarni

Contributed equally to this work with: Sharavari S. Kulkarni, John R. Spence, Christian J. Willenborg

Affiliation Department of Agricultural, Food, and Nutritional Science, University of Alberta, Edmonton, Alberta, Canada

Lloyd M. Dosdall

† Deceased.

Affiliation Department of Agricultural, Food, and Nutritional Science, University of Alberta, Edmonton, Alberta, Canada

John R. Spence

Contributed equally to this work with: Sharavari S. Kulkarni, John R. Spence, Christian J. Willenborg

Affiliation Department of Renewable Resources, University of Alberta, Edmonton, Alberta, Canada

Christian J. Willenborg

Contributed equally to this work with: Sharavari S. Kulkarni, John R. Spence, Christian J. Willenborg

chris.willenborg@usask.ca

Affiliation Department of Plant Sciences, College of Agriculture and Bioresources, University of Saskatchewan, Saskatoon, Saskatchewan, Canada

Competing Interests

We have the following interest: This study was funded in part by the Alberta Canola Producers Commission. There are no patents, products in development or marketed products to declare. This does not alter our adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceptualization: SSK LMD CJW. Formal analysis: CJW SSK JRS. Funding acquisition: CJW LMD. Investigation: SSK. Methodology: SSK LMD CJW. Resources: LMD CJW. Supervision: CJW LMD JRS. Validation: SSK CJW JRS. Writing – original draft: SSK CJW. Writing – review & editing: CJW JRS.