About the Authors

Xiaochen Cheng

Affiliation College of Marine Life Sciences, Ocean University of China, Qingdao, P. R. China

Ziyu Shao

Contributed equally to this work with: Ziyu Shao, Chengbo Li, Lejun Yu, Mazhar Ali Raja

Affiliation College of Marine Life Sciences, Ocean University of China, Qingdao, P. R. China

Chengbo Li

Contributed equally to this work with: Ziyu Shao, Chengbo Li, Lejun Yu, Mazhar Ali Raja

Affiliation College of Marine Life Sciences, Ocean University of China, Qingdao, P. R. China

Lejun Yu

Contributed equally to this work with: Ziyu Shao, Chengbo Li, Lejun Yu, Mazhar Ali Raja

Affiliation College of Marine Life Sciences, Ocean University of China, Qingdao, P. R. China

Mazhar Ali Raja

Contributed equally to this work with: Ziyu Shao, Chengbo Li, Lejun Yu, Mazhar Ali Raja

Affiliation College of Marine Life Sciences, Ocean University of China, Qingdao, P. R. China

Chenguang Liu

liucg@ouc.edu.cn

Affiliation College of Marine Life Sciences, Ocean University of China, Qingdao, P. R. China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: XCC CGL. Data curation: XCC ZYS. Formal analysis: XCC ZYS CBL. Investigation: XCC ZYS. Methodology: XCC ZYS LJY. Project administration: CGL. Resources: ZYS LJY. Software: XCC CBL. Supervision: CGL LJY. Validation: CGL LJY. Visualization: XCC CBL. Writing – original draft: XCC CBL MAR. Writing – review & editing: XCC MAR.