About the Authors

Masaya Tsuboi

Affiliation Laboratory of Veterinary Pathology, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan

Manabu Watanabe

Affiliation Laboratory of Functional Genomics, Graduate School of Frontier Sciences, The University of Tokyo, Tokyo, Japan

Kazumi Nibe

Affiliation Japan Animal Referral Medical Center, Kanagawa, Japan

Natsuko Yoshimi

Affiliation D&C Veterinary Clinic, Ibaraki, Japan

Akihisa Kato

Affiliation D&C Veterinary Clinic, Ibaraki, Japan

Masahiro Sakaguchi

Affiliation Laboratory of Veterinary Microbiology I, School of Veterinary Medicine, Azabu University, Kanagawa, Japan

Osamu Yamato

Affiliation Laboratory of Clinical Pathology, Joint Faculty of Veterinary Medicine, Kagoshima University, Kagoshima, Japan

Miyuu Tanaka

Affiliation Laboratory of Veterinary Pathology, Graduate School of Life and Environmental Sciences, Osaka Prefecture University, Osaka, Japan

Mitsuru Kuwamura

Affiliation Laboratory of Veterinary Pathology, Graduate School of Life and Environmental Sciences, Osaka Prefecture University, Osaka, Japan

Kazuya Kushida

Affiliation Laboratory of Clinical Pathology, Joint Faculty of Veterinary Medicine, Kagoshima University, Kagoshima, Japan

Takashi Ishikura

Affiliation Thermo Fisher Scientific, Life Technologies Japan Ltd., Tokyo, Japan

Tomoyuki Harada

Affiliation Laboratory of Veterinary Pathology, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan

James Kenn Chambers

Affiliation Laboratory of Veterinary Pathology, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan

Sumio Sugano

Affiliation Laboratory of Functional Genomics, Graduate School of Frontier Sciences, The University of Tokyo, Tokyo, Japan

Kazuyuki Uchida

auchidak@mail.ecc.u-tokyo.ac.jp

Affiliation Laboratory of Veterinary Pathology, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan

Hiroyuki Nakayama

Affiliation Laboratory of Veterinary Pathology, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan

Competing Interests

Although one of the authors (TI) is employed by a commercial company, (Thermo Fisher Scientific, Life Technologies Japan Ltd.), he has no competing interests concerning this work. This does not alter our adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceptualization: M. Tsuboi MW KU. Funding acquisition: M. Tsuboi. Investigation: M. Tsuboi MW TH. Project administration: KU. Resources: KN NY AK MS OY M. Tanaka MK. Software: TI. Supervision: SS HN. Validation: MS OY KK. Writing – original draft: M. Tsuboi. Writing – review & editing: MW KU JKC HN.