About the Authors

Ying Sun

Contributed equally to this work with: Ying Sun, Yu Huang, Guojun Hu

Affiliations State Key Laboratory of Biocontrol, Institute of Aquatic Economic Animals and Guangdong Provincial Key Laboratory for Aquatic Economic Animals, School of Life Sciences, Sun Yat-Sen University, Guangzhou, China, Shenzhen Key Lab of Marine Genomics, Guangdong Provincial Key Lab of Molecular Breeding in Marine Economic Animals, BGI, Shenzhen, China

Yu Huang

Contributed equally to this work with: Ying Sun, Yu Huang, Guojun Hu

Affiliation Shenzhen Key Lab of Marine Genomics, Guangdong Provincial Key Lab of Molecular Breeding in Marine Economic Animals, BGI, Shenzhen, China

Guojun Hu

Contributed equally to this work with: Ying Sun, Yu Huang, Guojun Hu

Affiliation Shenzhen Key Lab of Marine Genomics, Guangdong Provincial Key Lab of Molecular Breeding in Marine Economic Animals, BGI, Shenzhen, China

Xinhui Zhang

Affiliation Shenzhen Key Lab of Marine Genomics, Guangdong Provincial Key Lab of Molecular Breeding in Marine Economic Animals, BGI, Shenzhen, China

Zhiqiang Ruan

Affiliation Shenzhen Key Lab of Marine Genomics, Guangdong Provincial Key Lab of Molecular Breeding in Marine Economic Animals, BGI, Shenzhen, China

Xiaomeng Zhao

Affiliation Shenzhen Key Lab of Marine Genomics, Guangdong Provincial Key Lab of Molecular Breeding in Marine Economic Animals, BGI, Shenzhen, China

Chuanyu Guo

Affiliation Shenzhen Key Lab of Marine Genomics, Guangdong Provincial Key Lab of Molecular Breeding in Marine Economic Animals, BGI, Shenzhen, China

Zhujing Tang

Affiliation State Key Laboratory of Biocontrol, Institute of Aquatic Economic Animals and Guangdong Provincial Key Laboratory for Aquatic Economic Animals, School of Life Sciences, Sun Yat-Sen University, Guangzhou, China

Xiaofeng Li

Affiliation Shenzhen Key Lab of Marine Genomics, Guangdong Provincial Key Lab of Molecular Breeding in Marine Economic Animals, BGI, Shenzhen, China

Xinxin You

Affiliation Shenzhen Key Lab of Marine Genomics, Guangdong Provincial Key Lab of Molecular Breeding in Marine Economic Animals, BGI, Shenzhen, China

Haoran Lin

lsslhr@mail.sysu.edu.cn (Hl); lsszy@mail.sysu.edu.cn (YZ); shiqiong@genomics.cn (QS)

Affiliation State Key Laboratory of Biocontrol, Institute of Aquatic Economic Animals and Guangdong Provincial Key Laboratory for Aquatic Economic Animals, School of Life Sciences, Sun Yat-Sen University, Guangzhou, China

Yong Zhang

lsslhr@mail.sysu.edu.cn (Hl); lsszy@mail.sysu.edu.cn (YZ); shiqiong@genomics.cn (QS)

Affiliation State Key Laboratory of Biocontrol, Institute of Aquatic Economic Animals and Guangdong Provincial Key Laboratory for Aquatic Economic Animals, School of Life Sciences, Sun Yat-Sen University, Guangzhou, China

Qiong Shi

lsslhr@mail.sysu.edu.cn (Hl); lsszy@mail.sysu.edu.cn (YZ); shiqiong@genomics.cn (QS)

Affiliations Shenzhen Key Lab of Marine Genomics, Guangdong Provincial Key Lab of Molecular Breeding in Marine Economic Animals, BGI, Shenzhen, China, Center for Marine Research, School of Life Sciences and Oceanography, Shenzhen University, Shenzhen, China

Competing Interests

BGI-Shenzhen provided support in the form of salaries for authors YS, YH, GH, X. Zhang, ZR, X. Zhao, CG, XL, XY and QS. This does not alter our adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceptualization: YS HL YZ QS. Data curation: YH QS. Formal analysis: YH X. Zhang XL. Funding acquisition: QS YZ. Investigation: HL YZ ZR. Methodology: GH ZR X. Zhao YH. Project administration: GH. Resources: ZR X. Zhao YZ QS. Software: YH X. Zhang XY XL. Supervision: YS QS YZ. Validation: CG ZT ZR. Visualization: CG ZT FL. Writing – original draft: YS YH GH X. Zhang X. Zhao. Writing – review & editing: YH QS YZ.