About the Authors

Gunther Antonissen

Contributed equally to this work with: Gunther Antonissen, Roel Haesendonck

Gunther.Antonissen@UGent.be

Affiliations Department of Pharmacology, Toxicology and Biochemistry, Faculty of Veterinary Medicine, Ghent University, Merelbeke, Belgium, Department of Pathology, Bacteriology and Avian Diseases, Faculty of Veterinary Medicine, Ghent University, Merelbeke, Belgium

Roel Haesendonck

Contributed equally to this work with: Gunther Antonissen, Roel Haesendonck

Affiliation Department of Pathology, Bacteriology and Avian Diseases, Faculty of Veterinary Medicine, Ghent University, Merelbeke, Belgium

Mathias Devreese

Affiliation Department of Pharmacology, Toxicology and Biochemistry, Faculty of Veterinary Medicine, Ghent University, Merelbeke, Belgium

Nathan Broekaert

Affiliation Department of Pharmacology, Toxicology and Biochemistry, Faculty of Veterinary Medicine, Ghent University, Merelbeke, Belgium

Elin Verbrugghe

Affiliation Department of Pathology, Bacteriology and Avian Diseases, Faculty of Veterinary Medicine, Ghent University, Merelbeke, Belgium

Sarah De Saeger

Affiliation Department of Bioanalysis, Faculty of Pharmaceutical Sciences, Ghent University, Ghent, Belgium

Kris Audenaert

Affiliation Department of Applied Biosciences, Faculty of Bioscience Engineering, Ghent University, Ghent, Belgium

Freddy Haesebrouck

Affiliation Department of Pathology, Bacteriology and Avian Diseases, Faculty of Veterinary Medicine, Ghent University, Merelbeke, Belgium

Frank Pasmans

Affiliation Department of Pathology, Bacteriology and Avian Diseases, Faculty of Veterinary Medicine, Ghent University, Merelbeke, Belgium

Richard Ducatelle

Affiliation Department of Pathology, Bacteriology and Avian Diseases, Faculty of Veterinary Medicine, Ghent University, Merelbeke, Belgium

Siska Croubels

‡ These authors also contributed equally to this work.

Affiliation Department of Pharmacology, Toxicology and Biochemistry, Faculty of Veterinary Medicine, Ghent University, Merelbeke, Belgium

An Martel

‡ These authors also contributed equally to this work.

Affiliation Department of Pathology, Bacteriology and Avian Diseases, Faculty of Veterinary Medicine, Ghent University, Merelbeke, Belgium

Competing Interests

The authors have the following interests: This work received the financial support from Biomin GmbH, Herzogenburg, Austria. There are no patents, products in development or marketed products to declare. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceptualization: GA RH FP SC AM. Investigation: GA RH MD NB EV SDS RD. Resources: KA FH SC. Visualization: GA RH. Writing – original draft: GA RH NB. Writing – review & editing: MD EV SDS KA FH FP RD SC AM.