About the Authors

Adrián Arroyo

Affiliation Institute of Archaeology, University College London, 31–34 Gordon Square, London, WC1H 0PY, United Kingdom

Satoshi Hirata

hirata.satoshi.8z@kyoto-u.ac.jp (SH); i.torre@ucl.ac.uk (IdlT)

Affiliation Kumamoto Sanctuary, Wildlife Research Center, Kyoto University, 990 Ohtao, Misumi, Uki, Kumamoto, 869–201, Japan

Tetsuro Matsuzawa

Affiliations Kyoto University, Institute for Advanced Studies, Kyoto, Japan, Primate Research Institute, Kyoto University, Aichi, Japan, Japan Monkey Centre, Aichi, Japan

Ignacio de la Torre

hirata.satoshi.8z@kyoto-u.ac.jp (SH); i.torre@ucl.ac.uk (IdlT)

Affiliation Institute of Archaeology, University College London, 31–34 Gordon Square, London, WC1H 0PY, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: IdlT TM. Formal analysis: AA. Funding acquisition: SH TM IdlT. Investigation: SH. Methodology: SH IdlT. Writing – original draft: AA. Writing – review & editing: AA SH TM IdlT.