About the Authors

Min-Goo Seo

Affiliations College of Veterinary Medicine, Kyungpook National University, Daegu, South Korea, Animal and Plant Quarantine Agency, Gimcheon, South Korea

Seung-Hun Lee

Affiliation College of Veterinary Medicine, Kyungpook National University, Daegu, South Korea

In-Ohk Ouh

Affiliation Animal and Plant Quarantine Agency, Gimcheon, South Korea

Gwang Hyeop Lee

Affiliation Seowon Equine Clinic, Jeju, South Korea

Youn-Kyoung Goo

Affiliation Department of Parasitology and Tropical Medicine, Kyungpook National University School of Medicine, Daegu, South Korea

Seungjoon Kim

Affiliation College of Veterinary Medicine, Kyungpook National University, Daegu, South Korea

Oh-Deog Kwon

Affiliation College of Veterinary Medicine, Kyungpook National University, Daegu, South Korea

Dongmi Kwak

dmkwak@knu.ac.kr

Affiliations College of Veterinary Medicine, Kyungpook National University, Daegu, South Korea, Cardiovascular Research Institute, Kyungpook National University, Daegu, South Korea

Competing Interests

The authors have declared that no competing interests exist. Author GHL is employed by the Seowon Equine Clinic, which has links with commercial sponsors, however this does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceptualization: MGS DK. Data curation: MGS YKG SK ODK DK. Formal analysis: MGS SHL DK. Funding acquisition: DK. Investigation: MGS SHL DK. Methodology: MGS SHL DK. Project administration: SHL DK. Resources: MGS SHL IOO GHL ODK DK. Software: MGS SHL IOO. Supervision: DK. Validation: MGS SHL IOO DK. Visualization: MGS SHL DK. Writing – original draft: MGS SHL IOO DK. Writing – review & editing: MGS SHL YKG SK ODK DK.