About the Authors

Darci Rush

Contributed equally to this work with: Darci Rush, Kate A. Osborne

darci.rush@nioz.nl

Affiliation School of Civil Engineering & Geosciences, Newcastle University, Drummond Building, Newcastle upon Tyne, NE1 7RU, Newcastle-upon-Tyne, United Kingdom

Kate A. Osborne

Contributed equally to this work with: Darci Rush, Kate A. Osborne

Affiliation School of Civil Engineering & Geosciences, Newcastle University, Drummond Building, Newcastle upon Tyne, NE1 7RU, Newcastle-upon-Tyne, United Kingdom

Daniel Birgel

Affiliation Institute of Geology, University of Hamburg, Hamburg, Germany

Andreas Kappler

Affiliations Center for Applied Geoscience, University of Tübingen, Tübingen, Germany, Center for Geomicrobiology, Department of Bioscience, Ny Munkegade 116, 8000, Aarhus C, Denmark

Hisako Hirayama

Affiliation Department of Subsurface Geobiological Analysis and Research, Japan Agency for Marine-Earth Science & Technology (JAMSTEC), Yokosuka, Japan

Jörn Peckmann

Affiliations Institute of Geology, University of Hamburg, Hamburg, Germany, Department of Geodynamics and Sedimentology, University of Vienna, 1090, Vienna, Austria

Simon W. Poulton

Affiliation School of Earth and Environment, University of Leeds, Leeds, LS2 9JT, United Kingdom

Julia C. Nickel

Affiliation GFZ German Research Centre for Geosciences, Telegrafenberg, D-14473, Potsdam, Germany

Kai Mangelsdorf

Affiliation GFZ German Research Centre for Geosciences, Telegrafenberg, D-14473, Potsdam, Germany

Marina Kalyuzhnaya

Affiliation Faculty of Biology, San Diego State University, 5500 Campanile Drive, San Diego, 92182, United States of America

Frances R. Sidgwick

Affiliation School of Civil Engineering & Geosciences, Newcastle University, Drummond Building, Newcastle upon Tyne, NE1 7RU, Newcastle-upon-Tyne, United Kingdom

Helen M. Talbot

Affiliation School of Civil Engineering & Geosciences, Newcastle University, Drummond Building, Newcastle upon Tyne, NE1 7RU, Newcastle-upon-Tyne, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: KAO DR HMT. Formal analysis: DR KAO HMT. Investigation: KAO DR FRS. Methodology: KAO DR HMT. Resources: DB AK HH JP SWP JCN KM MK. Writing – original draft: DR KO HMT. Writing – review & editing: DB AK HH JP SWP JCN KM MK.