About the Authors

Kenneth Day

Contributed equally to this work with: Kenneth Day, Lindsay L. Waite

Affiliation HudsonAlpha Institute for Biotechnology, Huntsville, Alabama, United States of America

Lindsay L. Waite

Contributed equally to this work with: Kenneth Day, Lindsay L. Waite

Affiliations HudsonAlpha Institute for Biotechnology, Huntsville, Alabama, United States of America, University of Alabama at Birmingham, School of Public Health, Department of Biostatistics, Birmingham, Alabama, United States of America

Arnald Alonso

Current address: Rheumatology Research Group, Vall d'Hebron Hospital Research Institute, Barcelona, Spain

Affiliation HudsonAlpha Institute for Biotechnology, Huntsville, Alabama, United States of America

Marguerite R. Irvin

Affiliation University of Alabama at Birmingham, School of Public Health, Department of Epidemiology, Birmingham, Alabama, United States of America

Degui Zhi

Affiliation University of Alabama at Birmingham, School of Public Health, Department of Biostatistics, Birmingham, Alabama, United States of America

Krista S. Thibeault

Affiliation HudsonAlpha Institute for Biotechnology, Huntsville, Alabama, United States of America

Stella Aslibekyan

Affiliation University of Alabama at Birmingham, School of Public Health, Department of Epidemiology, Birmingham, Alabama, United States of America

Bertha Hidalgo

Affiliation University of Alabama at Birmingham, School of Public Health, Department of Biostatistics, Birmingham, Alabama, United States of America

Ingrid B. Borecki

Affiliation Washington University, Division of Statistical Genomics, St. Louis, Missouri, United States of America

Jose M. Ordovas

Affiliations Tufts University, Jean Mayer USDA Human Nutrition Research Center on Aging, Boston, Massachusetts, United States of America, IMDEA-Food, Madrid, Spain

Donna K. Arnett

Current address: University of Kentucky College of Public Health, Lexington, Kentucky, United States of America

Affiliation University of Alabama at Birmingham, School of Public Health, Department of Epidemiology, Birmingham, Alabama, United States of America

Hemant K. Tiwari

‡ These authors also contributed equally to this work.

Affiliation University of Alabama at Birmingham, School of Public Health, Department of Biostatistics, Birmingham, Alabama, United States of America

Devin M. Absher

dabsher@hudsonalpha.org

‡ These authors also contributed equally to this work.

Affiliation HudsonAlpha Institute for Biotechnology, Huntsville, Alabama, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: KD LLW AA HKT DMA. Data curation: MRI. Formal analysis: KD LLW AA MRI DZ SA BH IBB JMO DKA HKT DMA. Funding acquisition: DKA HKT DMA. Investigation: KD LLW KST DMA. Methodology: LLW HKT DMA AA. Project administration: DKA HKT DMA. Resources: MRI DZ SA BH IBB JMO DKA HKT DMA. Software: AA MRI LLW DZ. Supervision: DKA HKT DMA. Validation: SA BH. Visualization: KD LLW. Writing – original draft: KD LLW. Writing – review & editing: KD LLW HKT DMA.