About the Authors

Heba F. Alzan

Affiliations Department of Veterinary Microbiology and Pathology, College of Veterinary Medicine, Washington State University, Pullman, WA, United States of America, Parasitology and Animal Diseases Department, National Research Center, Dokki, Giza, Egypt

Audrey O. T. Lau

Contributed equally to this work with: Audrey O. T. Lau, Donald P. Knowles

Affiliation The National Institute of Allergy and Infectious Diseases, 5601 Fishers Lane, MSC 9823, Bethesda, MD, United States of America

Donald P. Knowles

Contributed equally to this work with: Audrey O. T. Lau, Donald P. Knowles

Affiliations Department of Veterinary Microbiology and Pathology, College of Veterinary Medicine, Washington State University, Pullman, WA, United States of America, Animal Disease Research Unit, United States Department of Agricultural—Agricultural Research Service, Pullman, WA, United States of America

David R. Herndon

Affiliation Animal Disease Research Unit, United States Department of Agricultural—Agricultural Research Service, Pullman, WA, United States of America

Massaro W. Ueti

Affiliations Department of Veterinary Microbiology and Pathology, College of Veterinary Medicine, Washington State University, Pullman, WA, United States of America, Animal Disease Research Unit, United States Department of Agricultural—Agricultural Research Service, Pullman, WA, United States of America

Glen A. Scoles

Affiliations Department of Veterinary Microbiology and Pathology, College of Veterinary Medicine, Washington State University, Pullman, WA, United States of America, Animal Disease Research Unit, United States Department of Agricultural—Agricultural Research Service, Pullman, WA, United States of America

Lowell S. Kappmeyer

Affiliation Animal Disease Research Unit, United States Department of Agricultural—Agricultural Research Service, Pullman, WA, United States of America

Carlos E. Suarez

ces@vetmed.wsu.edu

Affiliations Department of Veterinary Microbiology and Pathology, College of Veterinary Medicine, Washington State University, Pullman, WA, United States of America, Animal Disease Research Unit, United States Department of Agricultural—Agricultural Research Service, Pullman, WA, United States of America

Competing Interests

These authors have declared that no competing interest exist.

Author Contributions

Conceptualization: CES HFA. Data curation: HFA CES LSK DRH AOTL. Formal analysis: HFA. Funding acquisition: DPK CES HFA. Investigation: HFA CES. Methodology: HFA. Project administration: DPK CES. Resources: HFA CES MWU LSK DRH GAS. Software: HFA. Supervision: CES. Validation: HFA CES AOTL. Visualization: HFA CES. Writing – original draft: CES HFA. Writing – review & editing: CES HFA DPK AOTL LSK DRH MWU.