About the Authors

Radka Gromnicova

Affiliation Department of Life, Health and Chemical Sciences, The Open University, Milton Keynes, United Kingdom

Mehmet Kaya

Affiliation Department of Physiology, Koc University School of Medicine, Istanbul, Turkey

Ignacio A. Romero

Affiliation Department of Life, Health and Chemical Sciences, The Open University, Milton Keynes, United Kingdom

Phil Williams

Affiliation Midatech Pharma plc, Abingdon, United Kingdom

Simon Satchell

Affiliation School of Clinical Sciences, University of Bristol, Bristol, United Kingdom

Basil Sharrack

Affiliation Department of Neurology, University of Sheffield, Sheffield, United Kingdom

David Male

D.K.Male@Open.ac.uk

Affiliation Department of Life, Health and Chemical Sciences, The Open University, Milton Keynes, United Kingdom

Competing Interests

PW holds share options in Midatech Pharma plc. DM was previously on the Scientific advisory board of this company (until May 2016). This association does not alter our adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceptualization: DM MK IAR. Formal analysis: RG DM. Funding acquisition: BS DM. Investigation: RG MK PW. Methodology: RG DM MK. Project administration: DM. Resources: IAR SCS. Supervision: IAR BS. Validation: RG MK DM. Visualization: RG DM. Writing – original draft: RG DM. Writing – review & editing: RG MK IAR PW SCS BS DM.