About the Authors

Frauke Seehusen

Affiliation Department of Pathology, University of Veterinary Medicine, Hannover, Germany

Seham A. Al-Azreg

Affiliation Department of Pathology, University of Veterinary Medicine, Hannover, Germany

Barbara B. Raddatz

Affiliation Department of Pathology, University of Veterinary Medicine, Hannover, Germany

Verena Haist

Affiliations Department of Pathology, University of Veterinary Medicine, Hannover, Germany, Boehringer Ingelheim Veterinary Research Center GmbH & Co. KG, Hannover, Germany

Christina Puff

Affiliation Department of Pathology, University of Veterinary Medicine, Hannover, Germany

Ingo Spitzbarth

Affiliation Department of Pathology, University of Veterinary Medicine, Hannover, Germany

Reiner Ulrich

Affiliations Department of Pathology, University of Veterinary Medicine, Hannover, Germany, Friedrich-Loeffler-Institut, Federal Research Institute for Animal Health, Greifswald - Insel Riems, Germany

Wolfgang Baumgärtner

wolfgang.baumgaertner@tiho-hannover.de

Affiliation Department of Pathology, University of Veterinary Medicine, Hannover, Germany

Competing Interests

VH is employed by Boehringer Ingelheim Pharma GmbH&Co KG. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: FS VH CP RU WB. Performed the experiments: FS SA BBR IS CP. Analyzed the data: FS VH BBR IS CP. Contributed reagents/materials/analysis tools: SA CP RU WB. Wrote the paper: FS SA BBR CP IS WB.