About the Authors

Tsepo Lebiletsa Tsekoa

Affiliation Biosciences Unit, Council for Scientific and Industrial Research, Pretoria, South Africa

Therese Lotter-Stark

Affiliation Biosciences Unit, Council for Scientific and Industrial Research, Pretoria, South Africa

Sindisiwe Buthelezi

Affiliation Biosciences Unit, Council for Scientific and Industrial Research, Pretoria, South Africa

Ereck Chakauya

Affiliation Biosciences Unit, Council for Scientific and Industrial Research, Pretoria, South Africa

Stoyan H. Stoychev

Affiliation Biosciences Unit, Council for Scientific and Industrial Research, Pretoria, South Africa

Claude Sabeta

Affiliation ARC-Onderstepoort Veterinary Institute, Onderstepoort, South Africa

Wonderful Shumba

Affiliation ARC-Onderstepoort Veterinary Institute, Onderstepoort, South Africa

Baby Phahladira

Affiliation ARC-Onderstepoort Veterinary Institute, Onderstepoort, South Africa

Steve Hume

Affiliation Kentucky Bioprocessing, Owensboro, Kentucky, United States

Josh Morton

Affiliation Kentucky Bioprocessing, Owensboro, Kentucky, United States

Charles E. Rupprecht

Affiliation The Wistar Institute, Philadelphia, Pennsylvania, United States

Herta Steinkellner

Affiliation Department of Applied Genetics and Cell Biology, University of Natural Resources and Life Sciences, Vienna, Austria

Michael Pauly

Affiliation Mapp Biopharmaceutical, San Diego, California, United States

Larry Zeitlin

Affiliation Mapp Biopharmaceutical, San Diego, California, United States

Kevin Whaley

Affiliation Mapp Biopharmaceutical, San Diego, California, United States

Rachel Chikwamba

rchikwamba@csir.co.za

Affiliation Biosciences Unit, Council for Scientific and Industrial Research, Pretoria, South Africa

Competing Interests

LZ and KW are owners of Mapp Biopharmaceutical, a commercial company. MP is employed by Mapp Biopharmaceutical, while JM and SH are employed by Kentucky Bioprocessing, a commercial company. This does not alter the authors’ adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: RC LZ KW CS HS CR. Performed the experiments: TT TLS MP HS SS SB WS BP EC SH JM. Analyzed the data: EC TT HS SS CS MP SH JM. Contributed reagents/materials/analysis tools: CS HS CR. Wrote the paper: TT TLS SB EC SS CS WS BP SH JM CR HS MP LZ KW RC.