About the Authors

Keijiro Mizukami

Affiliation Section of Medical Genetics, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Karthik Raj

Affiliation Section of Medical Genetics, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Carl Osborne

Affiliation Veterinary Clinical Sciences, College of Veterinary Medicine, University of Minnesota, Saint Paul, MN, United States of America

Urs Giger

giger@vet.upenn.edu

Affiliation Section of Medical Genetics, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: UG KM. Performed the experiments: KM KR. Analyzed the data: UG KM. Contributed reagents/materials/analysis tools: UG CO KM. Wrote the paper: UG KM.