About the Authors

Andres H. Gutiérrez

Affiliation Institute for Immunology and Informatics, Department of Cell and Molecular Biology, University of Rhode Island, Providence, RI, United States of America

Crystal Loving

Affiliation Virus and Prion Diseases Research Unit, NADC, USDA ARS, Ames, IA, United States of America

Leonard Moise

Affiliations Institute for Immunology and Informatics, Department of Cell and Molecular Biology, University of Rhode Island, Providence, RI, United States of America, EpiVax Inc., Providence, RI, United States of America

Frances E. Terry

Affiliation EpiVax Inc., Providence, RI, United States of America

Susan L. Brockmeier

Affiliation Virus and Prion Diseases Research Unit, NADC, USDA ARS, Ames, IA, United States of America

Holly R. Hughes

Affiliation Virus and Prion Diseases Research Unit, NADC, USDA ARS, Ames, IA, United States of America

William D. Martin

Affiliation EpiVax Inc., Providence, RI, United States of America

Anne S. De Groot

dr.annie.degroot@gmail.com

Affiliations Institute for Immunology and Informatics, Department of Cell and Molecular Biology, University of Rhode Island, Providence, RI, United States of America, EpiVax Inc., Providence, RI, United States of America

Competing Interests

ADG and WDM are senior officers and majority shareholders at EpiVax, Inc., a privately-owned immunoinformatics and vaccine design company located in Providence, RI, USA. LM and FET are employees at EpiVax, in which LM holds stock options. ADG, WDM, LM and FET acknowledge that there is a potential conflict of interest related to their relationship with EpiVax and attest that the work contained in this research report is free of any bias that might be associated with the commercial goals of the company. This study was conducted in part with funding from National Pork Board. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials. There are no patents, products in development or marketed products to declare.

Author Contributions

Conceived and designed the experiments: AHG CL LM ADG WDM. Performed the experiments: AHG CL SLB HRH. Analyzed the data: AHG CL LM ADG FET. Contributed reagents/materials/analysis tools: AHG CL WDM ADG. Wrote the paper: AHG CL LM ADG WDM FET SLB HRH.