About the Authors

R. Hoare

rh30@stir.ac.uk

Affiliation Institute of Aquaculture, School of Natural Sciences, University of Stirling, Stirling, FK9 4LA, United Kingdom

K. D. Thompson

Affiliations Institute of Aquaculture, School of Natural Sciences, University of Stirling, Stirling, FK9 4LA, United Kingdom, Moredun Research Institute, Pentlands Science Park, Penicuik EH26 0PZ, United Kingdom

T. Herath

Affiliation Department of Animal Production, Welfare and Veterinary Sciences, Harper Adams University, Newport, Shropshire TF10 8NB, United Kingdom

B. Collet

Affiliation Marine Scotland, Marine Laboratory, 375 Victoria Road, Aberdeen AB11 9PA, United Kingdom

J. E. Bron

Affiliation Institute of Aquaculture, School of Natural Sciences, University of Stirling, Stirling, FK9 4LA, United Kingdom

A. Adams

Affiliation Institute of Aquaculture, School of Natural Sciences, University of Stirling, Stirling, FK9 4LA, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BC KDT AA. Performed the experiments: RH KDT. Analyzed the data: RH. Contributed reagents/materials/analysis tools: BC KDT. Wrote the paper: RH KDT TH AA. Performed the confocal microscopy: JEB. Advised on development of qPCR: TH.