About the Authors

Ayako Aoki-Yoshida

Affiliations Department of Applied Biological Chemistry, The University of Tokyo, Yayoi, Bunkyo-ku, Tokyo, Japan, National Institute of Livestock and Grassland Science, National Agriculture and Food Research Organization, Tsukuba, Ibaraki, Japan

Kiyoshi Yamada

Affiliation Department of Applied Biological Chemistry, The University of Tokyo, Yayoi, Bunkyo-ku, Tokyo, Japan

Satoshi Hachimura

Affiliation Research Center for Food Safety, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan

Toshihiro Sashihara

Affiliation Division of Research and Development, Meiji Co. Ltd., Odawara, Kanagawa, Japan

Shuji Ikegami

Affiliation Division of Research and Development, Meiji Co. Ltd., Odawara, Kanagawa, Japan

Makoto Shimizu

Affiliation Department of Applied Biological Chemistry, The University of Tokyo, Yayoi, Bunkyo-ku, Tokyo, Japan

Mamoru Totsuka

atotuka@mail.ecc.u-tokyo.ac.jp

Affiliation Department of Applied Biological Chemistry, The University of Tokyo, Yayoi, Bunkyo-ku, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AA MT. Performed the experiments: AA KY SH TS SI MS MT. Analyzed the data: AA. Contributed reagents/materials/analysis tools: AA KY SH TS SI MS MT. Wrote the paper: AA MT.