About the Authors

Irene Cano

Affiliation Aquatic Animal Disease, Centre for Environment, Fisheries and Aquaculture Science, Barrack Road, The Nothe, Weymouth, Dorset DT4 8UB, United Kingdom

David W. Verner-Jeffreys

david.verner-jeffreys@cefas.co.uk

Affiliation Aquatic Animal Disease, Centre for Environment, Fisheries and Aquaculture Science, Barrack Road, The Nothe, Weymouth, Dorset DT4 8UB, United Kingdom

Ronny van Aerle

Affiliation Aquatic Animal Disease, Centre for Environment, Fisheries and Aquaculture Science, Barrack Road, The Nothe, Weymouth, Dorset DT4 8UB, United Kingdom

Richard K. Paley

Affiliation Aquatic Animal Disease, Centre for Environment, Fisheries and Aquaculture Science, Barrack Road, The Nothe, Weymouth, Dorset DT4 8UB, United Kingdom

Edmund J. Peeler

Affiliation Aquatic Animal Disease, Centre for Environment, Fisheries and Aquaculture Science, Barrack Road, The Nothe, Weymouth, Dorset DT4 8UB, United Kingdom

Matthew Green

Affiliation Aquatic Animal Disease, Centre for Environment, Fisheries and Aquaculture Science, Barrack Road, The Nothe, Weymouth, Dorset DT4 8UB, United Kingdom

Georgina S. E. Rimmer

Affiliation Aquatic Animal Disease, Centre for Environment, Fisheries and Aquaculture Science, Barrack Road, The Nothe, Weymouth, Dorset DT4 8UB, United Kingdom

Jacqueline Savage

Affiliation Aquatic Animal Disease, Centre for Environment, Fisheries and Aquaculture Science, Barrack Road, The Nothe, Weymouth, Dorset DT4 8UB, United Kingdom

Claire L. Joiner

Affiliation Aquatic Animal Disease, Centre for Environment, Fisheries and Aquaculture Science, Barrack Road, The Nothe, Weymouth, Dorset DT4 8UB, United Kingdom

Amanda E. Bayley

Affiliation Aquatic Animal Disease, Centre for Environment, Fisheries and Aquaculture Science, Barrack Road, The Nothe, Weymouth, Dorset DT4 8UB, United Kingdom

Jason Mewett

Affiliation Aquatic Animal Disease, Centre for Environment, Fisheries and Aquaculture Science, Barrack Road, The Nothe, Weymouth, Dorset DT4 8UB, United Kingdom

Jonathan Hulland

Affiliation Aquatic Animal Disease, Centre for Environment, Fisheries and Aquaculture Science, Barrack Road, The Nothe, Weymouth, Dorset DT4 8UB, United Kingdom

Stephen W. Feist

Affiliation Aquatic Animal Disease, Centre for Environment, Fisheries and Aquaculture Science, Barrack Road, The Nothe, Weymouth, Dorset DT4 8UB, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DWVJ IC SWF RKP EJP. Performed the experiments: IC DWVJ MG CLJ AEB GSER SWF RKP JS JM JH. Analyzed the data: IC DWVJ SWF RvA. Wrote the paper: DWVJ IC SWF RvA RKP EJP.