About the Authors

Jian Zhang

Contributed equally to this work with: Jian Zhang, Huwei Yuan

Affiliations Center for Computational Biology, College of Biological Sciences and Technology, Beijing Forestry University, Beijing, China, Jiangsu Riverine Institute of Agricultural Sciences, Rugao, Jiangsu, China

Huwei Yuan

Contributed equally to this work with: Jian Zhang, Huwei Yuan

Affiliation National Engineering Laboratory for Forest Tree Breeding, Key Laboratory for Genetics and Breeding of Forest Trees and Ornamental Plants of Ministry of Education, College of Biological Sciences and Technology, Beijing Forestry University, Beijing, China

Min Li

Affiliation Jiangsu Riverine Institute of Agricultural Sciences, Rugao, Jiangsu, China

Yujuan Li

Affiliation Jiangsu Riverine Institute of Agricultural Sciences, Rugao, Jiangsu, China

Ying Wang

Affiliation Jiangsu Riverine Institute of Agricultural Sciences, Rugao, Jiangsu, China

Xiangjian Ma

Affiliation Jiangsu Riverine Institute of Agricultural Sciences, Rugao, Jiangsu, China

Yuan Zhang

Affiliation Jiangsu Riverine Institute of Agricultural Sciences, Rugao, Jiangsu, China

Feng Tan

Affiliation Jiangsu Riverine Institute of Agricultural Sciences, Rugao, Jiangsu, China

Rongling Wu

rwu@bjfu.edu.cn; rwu@phs.psu.edu

Affiliations Center for Computational Biology, College of Biological Sciences and Technology, Beijing Forestry University, Beijing, China, Center for Statistical Genetics, Pennsylvania State University, Hershey, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RW JZ. Performed the experiments: JZ HY ML YZ. Analyzed the data: HY YW. Contributed reagents/materials/analysis tools: YL XM FT. Wrote the paper: RW JZ HY.