About the Authors

Koshi Kitamura

Affiliation Graduate School of Science and Technology, Shizuoka University, Hamamatsu, Shizuoka, Japan

Satoshi Kakishima

kakishim@gmail.com

Affiliation Graduate School of Science and Technology, Shizuoka University, Hamamatsu, Shizuoka, Japan

Takashi Uehara

Affiliation Graduate School of Science and Technology, Shizuoka University, Hamamatsu, Shizuoka, Japan

Satoru Morita

Affiliation Department of Mathematical and Systems Engineering, Shizuoka University, Hamamatsu, Shizuoka, Japan

Kei-ichi Tainaka

Affiliation Graduate School of Science and Technology, Shizuoka University, Hamamatsu, Shizuoka, Japan

Jin Yoshimura

Affiliations Graduate School of Science and Technology, Shizuoka University, Hamamatsu, Shizuoka, Japan, Marine Biosystems Research Center, Chiba University, Kamogawa, Chiba, Japan, Department of Environmental and Forest Biology, State University of New York College of Environmental Science and Forestry, Syracuse, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KK SK KT JY. Performed the experiments: KK. Analyzed the data: KK SK TU SM. Wrote the paper: KK SK TU SM KT JY.