About the Authors

Linnea M. Carver

Affiliation Department of Biology, Sewanee: The University of the South, Sewanee, Tennessee, United States of America

Patricia Perlaky

Affiliation Raccoon Mountain Caverns, Chattanooga, Tennessee, United States of America

Alan Cressler

Affiliation Independent Researcher, Atlanta, Georgia, United States of America

Kirk S. Zigler

kzigler@sewanee.edu

Affiliation Department of Biology, Sewanee: The University of the South, Sewanee, Tennessee, United States of America

Competing Interests

The authors have declared that no competing interests exist. Raccoon Mountain Caverns provided support in the form of salary for one author [PP], but this does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: LMC KSZ. Performed the experiments: LMC PP AC KSZ. Analyzed the data: LMC KSZ. Contributed reagents/materials/analysis tools: LMC PP AC KSZ. Wrote the paper: LMC PP AC KSZ.