About the Authors

Arpron Leesombun

Affiliation National Research Center for Protozoan Diseases, Obihiro University of Agriculture and Veterinary Medicine, Inada-cho, Obihiro, Hokkaido 080–8555, Japan

Sookruetai Boonmasawai

Affiliation Department of Preclinic and Applied Animal Science, Faculty of Veterinary Science, Mahidol University, 999 Phutthamonthon Sai 4 Road Salaya, Phutthamonthon Nakhonpathom 73170, Thailand

Naomi Shimoda

Affiliation National Research Center for Protozoan Diseases, Obihiro University of Agriculture and Veterinary Medicine, Inada-cho, Obihiro, Hokkaido 080–8555, Japan

Yoshifumi Nishikawa

nisikawa@obihiro.ac.jp

Affiliation National Research Center for Protozoan Diseases, Obihiro University of Agriculture and Veterinary Medicine, Inada-cho, Obihiro, Hokkaido 080–8555, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AL YN. Performed the experiments: AL SB NS YN. Analyzed the data: AL YN. Contributed reagents/materials/analysis tools: AL SB NS. Wrote the paper: AL YN.