About the Authors

Dan Luo

Affiliation Department of Biochemistry & Molecular Biology, School of Life Sciences, Sichuan University, Chengdu, China

Caixia Wen

Affiliation Department of Biochemistry & Molecular Biology, School of Life Sciences, Sichuan University, Chengdu, China

Rongchuan Zhao

Affiliation Department of Biochemistry & Molecular Biology, School of Life Sciences, Sichuan University, Chengdu, China

Xinyu Liu

Affiliation Department of Biochemistry & Molecular Biology, School of Life Sciences, Sichuan University, Chengdu, China

Xinxin Liu

Affiliation Department of Biochemistry & Molecular Biology, School of Life Sciences, Sichuan University, Chengdu, China

Jingjing Cui

Affiliation Department of Biochemistry & Molecular Biology, School of Life Sciences, Sichuan University, Chengdu, China

Joshua G. Liang

Affiliation Clover Biopharmaceuticals, Chengdu, China

Peng Liang

liangpeng@scu.edu.cn; liang.peng@cloverbiopharma.com

Affiliations Department of Biochemistry & Molecular Biology, School of Life Sciences, Sichuan University, Chengdu, China, Clover Biopharmaceuticals, Chengdu, China, GenHunter Corporation, Grassmere Park, Nashville, United States of America

Competing Interests

These authors (PL and JGL) are affiliated with commercial companies: Clover Biopharmaceuticals and GenHunter Corporation. On behalf of all authors, the corresponding author declares that commercial affiliations do not alter the adherence to PLOS ONE policies on sharing data.

Author Contributions

Conceived and designed the experiments: PL JGL DL. Performed the experiments: DL CXW RCZ. Analyzed the data: PL DL. Contributed reagents/materials/analysis tools: XYL XXL JJC. Wrote the paper: PL JGL DL.