About the Authors

Maja Coulthard Hampson

Affiliations Bristol Zoological Society, Bristol, United Kingdom, School of Veterinary Sciences, University of Bristol, Bristol, United Kingdom

Christoph Schwitzer

cschwitzer@bristolzoo.org.uk

Affiliation Bristol Zoological Society, Bristol, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CS MCH. Performed the experiments: MCH. Analyzed the data: MCH. Wrote the paper: MCH CS.