About the Authors

Jae-kyung Kim

Affiliations The Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, 222 Banpo-daero, Seocho-gu, Seoul, 137–701, South Korea, Laboratory of Immune Network, Catholic Research Institute of Medical Science, The Catholic University of Korea, 222 Banpo-daero, Seocho-gu, Seoul, 137–701, South Korea

Seung Hoon Lee

Affiliations The Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, 222 Banpo-daero, Seocho-gu, Seoul, 137–701, South Korea, Laboratory of Immune Network, Catholic Research Institute of Medical Science, The Catholic University of Korea, 222 Banpo-daero, Seocho-gu, Seoul, 137–701, South Korea

Seon-Young Lee

Affiliations The Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, 222 Banpo-daero, Seocho-gu, Seoul, 137–701, South Korea, Laboratory of Immune Network, Catholic Research Institute of Medical Science, The Catholic University of Korea, 222 Banpo-daero, Seocho-gu, Seoul, 137–701, South Korea

Eun-Kyung Kim

Affiliations The Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, 222 Banpo-daero, Seocho-gu, Seoul, 137–701, South Korea, Laboratory of Immune Network, Catholic Research Institute of Medical Science, The Catholic University of Korea, 222 Banpo-daero, Seocho-gu, Seoul, 137–701, South Korea

Jeong-Eun Kwon

Affiliations The Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, 222 Banpo-daero, Seocho-gu, Seoul, 137–701, South Korea, Laboratory of Immune Network, Catholic Research Institute of Medical Science, The Catholic University of Korea, 222 Banpo-daero, Seocho-gu, Seoul, 137–701, South Korea

Hyeon-Beom Seo

Affiliations The Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, 222 Banpo-daero, Seocho-gu, Seoul, 137–701, South Korea, Laboratory of Immune Network, Catholic Research Institute of Medical Science, The Catholic University of Korea, 222 Banpo-daero, Seocho-gu, Seoul, 137–701, South Korea

Han Hee Lee

Affiliation Division of Gastroenterlogy, Department of Internal Medicine, College of Medicine, The Catholic University of Korea, Seoul, Republic of Korea

Bo-In Lee

Affiliation Division of Gastroenterlogy, Department of Internal Medicine, College of Medicine, The Catholic University of Korea, Seoul, Republic of Korea

Sung-Hwan Park

Affiliations The Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, 222 Banpo-daero, Seocho-gu, Seoul, 137–701, South Korea, Division of Rheumatology, Department of Internal Medicine, The Catholic University of Korea, Seoul, 137–040, South Korea

Mi-La Cho

iammila@catholic.ac.kr

Affiliations The Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, 222 Banpo-daero, Seocho-gu, Seoul, 137–701, South Korea, Laboratory of Immune Network, Catholic Research Institute of Medical Science, The Catholic University of Korea, 222 Banpo-daero, Seocho-gu, Seoul, 137–701, South Korea

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JKK MLC. Performed the experiments: JKK EKK HBS. Analyzed the data: JKK SYL EKK JEK HHL BIL SHP. Contributed reagents/materials/analysis tools: MLC. Wrote the paper: JKK SHL.