About the Authors

Emily Weiss

Affiliation Usher Institute for Population Health Sciences and Informatics, University of Edinburgh, Edinburgh, Scotland

Lina Zgaga

Affiliation Public Health and Primary Care, Trinity College Centre for Health Sciences, Dublin, Ireland

Stephanie Read

Affiliation Usher Institute for Population Health Sciences and Informatics, University of Edinburgh, Edinburgh, Scotland

Sarah Wild

Affiliation Usher Institute for Population Health Sciences and Informatics, University of Edinburgh, Edinburgh, Scotland

Malcolm G. Dunlop

Affiliation MRC Human Genetics Unit, MRC Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, Scotland

Harry Campbell

Affiliation Usher Institute for Population Health Sciences and Informatics, University of Edinburgh, Edinburgh, Scotland

Ruth McQuillan

Contributed equally to this work with: Ruth McQuillan, James F. Wilson

Affiliation Usher Institute for Population Health Sciences and Informatics, University of Edinburgh, Edinburgh, Scotland

James F. Wilson

Contributed equally to this work with: Ruth McQuillan, James F. Wilson

jim.wilson@ed.ac.uk

Affiliations Usher Institute for Population Health Sciences and Informatics, University of Edinburgh, Edinburgh, Scotland, MRC Human Genetics Unit, MRC Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, Scotland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EW JFW RM. Performed the experiments: EW. Analyzed the data: EW LZ SR. Contributed reagents/materials/analysis tools: HC MGD JFW. Wrote the paper: EW LZ SR SW HC MGD RM JFW.