About the Authors

Nitipong Homwong

Affiliations Department of Veterinary Population Medicine, College of Veterinary Medicine, University of Minnesota, Saint Paul, Minnesota, United States of America, Department of Animal Science, Kasetsart University, Kamphaeng Saen Campus, Kamphaeng Saen, Nakhon Pathom, Thailand

Andres Diaz

Affiliation Department of Veterinary Population Medicine, College of Veterinary Medicine, University of Minnesota, Saint Paul, Minnesota, United States of America

Stephanie Rossow

Affiliation Department of Veterinary Population Medicine, College of Veterinary Medicine, University of Minnesota, Saint Paul, Minnesota, United States of America

Max Ciarlet

Affiliation Vaccines Clinical Research and Development, GlaxoSmithKline Vaccines, Cambridge, Massachusetts, United States of America

Douglas Marthaler

marth027@umn.edu

Affiliation Department of Veterinary Population Medicine, College of Veterinary Medicine, University of Minnesota, Saint Paul, Minnesota, United States of America

Competing Interests

The authors declare they have no conflicts of interest, and the author affiliation with GlaxoSmithKline Vaccines does not alter the author's adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: NH DM. Performed the experiments: NH DM. Analyzed the data: NH AD. Contributed reagents/materials/analysis tools: NH SR DM. Wrote the paper: NH AD SR MC DM.