About the Authors

Yi-Fan Hong

Contributed equally to this work with: Yi-Fan Hong, Hangeun Kim

Affiliations Graduate School of Biotechnology and Institute of Life Science and Resources, Kyung Hee University, Yongin, 446–701, Republic of Korea, Skin Biotechnology Center, Kyung Hee University, Yongin, 446–701, Republic of Korea

Hangeun Kim

Contributed equally to this work with: Yi-Fan Hong, Hangeun Kim

Affiliations Graduate School of Biotechnology and Institute of Life Science and Resources, Kyung Hee University, Yongin, 446–701, Republic of Korea, Skin Biotechnology Center, Kyung Hee University, Yongin, 446–701, Republic of Korea

Hye Sun Kim

Current address: Department of Neurosurgery, Ajou University School of Medicine, Suwon, 443–749, Republic of Korea

Affiliation Graduate School of Biotechnology and Institute of Life Science and Resources, Kyung Hee University, Yongin, 446–701, Republic of Korea

Woo Jung Park

Current address: Department of Marine Food Science and Technology, Gangneung-Wonju National University, Gangneung, 210–702, Republic of Korea

Affiliation Graduate School of Biotechnology and Institute of Life Science and Resources, Kyung Hee University, Yongin, 446–701, Republic of Korea

Joo-Yun Kim

99djawl@hanmail.net (JYK); dkchung@khu.ac.kr (DKC)

Affiliation Graduate School of Biotechnology and Institute of Life Science and Resources, Kyung Hee University, Yongin, 446–701, Republic of Korea

Dae Kyun Chung

99djawl@hanmail.net (JYK); dkchung@khu.ac.kr (DKC)

Affiliations Graduate School of Biotechnology and Institute of Life Science and Resources, Kyung Hee University, Yongin, 446–701, Republic of Korea, Skin Biotechnology Center, Kyung Hee University, Yongin, 446–701, Republic of Korea, RNA Inc., #308 Life Sceince Building, Kyung Hee University Global Campus, Yongin, 446–701, Republic of Korea

Competing Interests

Dae Kyun Chung is employed by RNA Inc. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: HK DKC JYK. Performed the experiments: YFH HSK WJP HK JYK. Analyzed the data: HK DKC JYK. Contributed reagents/materials/analysis tools: HSK WJP. Wrote the paper: JYK HK YFH.