About the Authors

Izumi Matsudaira

Affiliation Department of Nuclear Medicine & Radiology, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan

Susumu Yokota

yokota.e1@tohoku.ac.jp

Affiliation Division of Developmental Cognitive Neuroscience, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan

Teruo Hashimoto

Affiliation Division of Developmental Cognitive Neuroscience, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan

Hikaru Takeuchi

Affiliation Division of Developmental Cognitive Neuroscience, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan

Kohei Asano

Affiliation Division of Developmental Cognitive Neuroscience, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan

Michiko Asano

Affiliation Division of Developmental Cognitive Neuroscience, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan

Yuko Sassa

Affiliation Division of Developmental Cognitive Neuroscience, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan

Yasuyuki Taki

Affiliations Department of Nuclear Medicine & Radiology, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan, Division of Developmental Cognitive Neuroscience, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan, Division of Medical Neuroimaging Analysis, Department of Community Medical Supports, Tohoku Medical Megabank Organization, Tohoku University, Sendai, Japan

Ryuta Kawashima

Affiliations Division of Developmental Cognitive Neuroscience, Institute of Development, Aging and Cancer, Tohoku University, Sendai, Japan, Department of Functional Brain Imaging, Institute of Development, Aging, and Cancer, Tohoku University, Sendai, Japan, Smart Aging International Research Center, Institute of Development, Aging, and Cancer, Tohoku University, Sendai, Japan

Competing Interests

This study received funding from the Kumon Institute of Education Co., Ltd. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: YT RK HT KA MA YS. Performed the experiments: SY HT KA MA YS YT. Analyzed the data: IM HT SY TH. Contributed reagents/materials/analysis tools: YT RK HT KA MA YS. Wrote the paper: IM SY YT.