About the Authors

Ningning Ma

Contributed equally to this work with: Ningning Ma, Lili Zhang

Affiliation Soil Ecology and Agro-ecological Engineering Research Center, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110164, People`s Republic of China

Lili Zhang

Contributed equally to this work with: Ningning Ma, Lili Zhang

llzhang@iae.ac.cn (LZ); ygh006@iae.ac.cn (GY)

Affiliation Soil Ecology and Agro-ecological Engineering Research Center, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110164, People`s Republic of China

Yulan Zhang

Affiliation Soil Ecology and Agro-ecological Engineering Research Center, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110164, People`s Republic of China

Lijie Yang

Affiliations Soil Ecology and Agro-ecological Engineering Research Center, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110164, People`s Republic of China, Graduate School, University of Chinese Academy of Sciences, Beijing 100049, People`s Republic of China

Chunxiao Yu

Affiliations Soil Ecology and Agro-ecological Engineering Research Center, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110164, People`s Republic of China, Graduate School, University of Chinese Academy of Sciences, Beijing 100049, People`s Republic of China

Guanghua Yin

llzhang@iae.ac.cn (LZ); ygh006@iae.ac.cn (GY)

Affiliation Soil Ecology and Agro-ecological Engineering Research Center, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110164, People`s Republic of China

Timothy A. Doane

Affiliation Department of Land, Air, and Water Resources, University of California Davis, Davis 95616, United State of America

Zhijie Wu

Affiliation Soil Ecology and Agro-ecological Engineering Research Center, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110164, People`s Republic of China

Ping Zhu

Affiliation Institute of Agricultural Resources and Environment, Jilin Academy of Agricultural Sciences, Changchun 130033, People`s Republic of China

Xingzhu Ma

Affiliation Institute of Soil Fertilizer and Environmental Resources, Heilongjiang Academy of Agricultural Sciences, Harbin, 150086, People`s Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NM LZ LY CY GY. Performed the experiments: NM LZ YZ PZ XM. Analyzed the data: NM LZ TAD ZW. Contributed reagents/materials/analysis tools: NM LZ YZ LY CY GY TAD ZW PZ XM. Wrote the paper: NM LZ LY CY GY TAD ZW PZ XM.