About the Authors

Yuelin Liu

Affiliations Hokkaido Agriculture Research Center, National Agriculture and Food Research Organization, Sapporo, Japan, Graduate School of Agriculture, Hokkaido University, Sapporo, Japan

Daisuke Tabata

Affiliation Graduate School of Agriculture, Hokkaido University, Sapporo, Japan

Ryozo Imai

rzi@affrc.go.jp

Affiliations Hokkaido Agriculture Research Center, National Agriculture and Food Research Organization, Sapporo, Japan, Graduate School of Agriculture, Hokkaido University, Sapporo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YL RI. Performed the experiments: YL DT. Analyzed the data: YL RI. Contributed reagents/materials/analysis tools: YL DT RI. Wrote the paper: YL RI.