About the Authors

Zhihui Chang

Contributed equally to this work with: Zhihui Chang, Yang Liu

changzh@bjfu.edu.cn (ZC); xuzhang@vt.edu (XZ)

Affiliation College of Forestry, Beijing Forestry University, Beijing, 100083, P.R. China

Yang Liu

Contributed equally to this work with: Zhihui Chang, Yang Liu

Affiliation College of Forestry, Beijing Forestry University, Beijing, 100083, P.R. China

Hui Dong

Affiliation College of Forestry, Beijing Forestry University, Beijing, 100083, P.R. China

Ke Teng

Affiliation College of Forestry, Beijing Forestry University, Beijing, 100083, P.R. China

Liebao Han

Affiliation College of Forestry, Beijing Forestry University, Beijing, 100083, P.R. China

Xunzhong Zhang

changzh@bjfu.edu.cn (ZC); xuzhang@vt.edu (XZ)

Affiliation Department of Crop and Soil Environmental Sciences, Virginia Polytechnic Institute and State University, Blacksburg, Virginia, 24061, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: XZ ZC YL. Performed the experiments: ZC YL HD. Analyzed the data: ZC YL HD. Contributed reagents/materials/analysis tools: ZC. Wrote the paper: ZC YL XZ HD KT LH.