About the Authors

Sandra M. Davern

Affiliation Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America

Timothy E. McKnight

Affiliation Electrical & Electronics Systems Research Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America

Robert F. Standaert

standaertrf@ornl.gov

Affiliations Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America, Biology & Soft Matter Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America, Department of Biochemistry and Cellular & Molecular Biology, University of Tennessee, Knoxville, Tennessee, United States of America

Jennifer L. Morrell-Falvey

Affiliations Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America, Department of Biochemistry and Cellular & Molecular Biology, University of Tennessee, Knoxville, Tennessee, United States of America

Elena D. Shpak

Affiliation Department of Biochemistry and Cellular & Molecular Biology, University of Tennessee, Knoxville, Tennessee, United States of America

Udaya C. Kalluri

Affiliation Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America

Joanna Jelenska

Affiliation Department of Molecular Genetics and Cell Biology, The University of Chicago, Chicago, Illinois, United States of America

Jean T. Greenberg

Affiliation Department of Molecular Genetics and Cell Biology, The University of Chicago, Chicago, Illinois, United States of America

Saed Mirzadeh

Affiliation Nuclear Security & Isotope Technology Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SMD TEM RFS UCK JJ JTG SM. Performed the experiments: SMD TEM JLM-F EDS UCK. Analyzed the data: SMD TEM RFS JJ JTG. Wrote the paper: SMD TEM RFS JJ JTG SM.