About the Authors

Kim E. Reuter

kimeleanorreuter@gmail.com

Affiliation Temple University, Department of Biology, Philadelphia, Pennsylvania, United States of America

Abigail R. Wills

Affiliation Mpingo Conservation & Development Initiative, Kilwa Masoko, Tanzania

Raymond W. Lee

Affiliation Washington State University, School of Biological Sciences, Pullman, Washington, United States of America

Erik E. Cordes

Affiliation Temple University, Department of Biology, Philadelphia, Pennsylvania, United States of America

Brent J. Sewall

Affiliation Temple University, Department of Biology, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KER EEC BJS ARW RWL. Performed the experiments: KER ARW RWL. Analyzed the data: KER RWL EEC BJS. Contributed reagents/materials/analysis tools: RWL EEC. Wrote the paper: KER ARW RWL EEC BJS.