About the Authors

Shuichi Fukuyoshi

Contributed equally to this work with: Shuichi Fukuyoshi, Masaharu Kometani

Affiliation Institute of Medical, Pharmaceutical and Health Sciences, Kanazawa University, Kakuma-machi, Kanazawa, Ishikawa, 920–1192, Japan

Masaharu Kometani

Contributed equally to this work with: Shuichi Fukuyoshi, Masaharu Kometani

Affiliation Institute of Medical, Pharmaceutical and Health Sciences, Kanazawa University, Kakuma-machi, Kanazawa, Ishikawa, 920–1192, Japan

Yurie Watanabe

Affiliation Institute of Medical, Pharmaceutical and Health Sciences, Kanazawa University, Kakuma-machi, Kanazawa, Ishikawa, 920–1192, Japan

Masahiro Hiratsuka

Affiliation Graduate School of Pharmaceutical Sciences, Tohoku University, 6–3 Aoba, Aramaki, Aoba-ku, Sendai, 980–8578, Japan

Noriyuki Yamaotsu

Affiliation School of Pharmacy, Kitasato University, 5-9-1 Shirokane, Minato-ku, Tokyo, 108–8641, Japan

Shuichi Hirono

Affiliation School of Pharmacy, Kitasato University, 5-9-1 Shirokane, Minato-ku, Tokyo, 108–8641, Japan

Noriyoshi Manabe

Affiliation Faculty of Pharmaceutical Sciences, Tohoku Pharmaceutical University, 4-4-1 Komatsushima, Aoba-ku, Sendai, Miyagi, 981–8558, Japan

Ohgi Takahashi

Affiliation Faculty of Pharmaceutical Sciences, Tohoku Pharmaceutical University, 4-4-1 Komatsushima, Aoba-ku, Sendai, Miyagi, 981–8558, Japan

Akifumi Oda

oda@p.kanazawa-u.ac.jp

Affiliations Institute of Medical, Pharmaceutical and Health Sciences, Kanazawa University, Kakuma-machi, Kanazawa, Ishikawa, 920–1192, Japan, Institute for Protein Research, Osaka University, 3–2 Yamadaoka, Suita, Osaka, 565–0871, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SF YW OT AO. Performed the experiments: SF MK YW NM AO. Analyzed the data: MK MH NY SH AO. Contributed reagents/materials/analysis tools: MH NY SH. Wrote the paper: SF MK AO.