About the Authors

Hiroyuki Tada

htada@dent.tohoku.ac.jp

Affiliations Department of Oral Disease Research, National Center for Geriatrics and Gerontology, Obu, Aichi, Japan, Division of Oral Microbiology, Tohoku University Graduate School of Dentistry, Sendai, Miyagi, Japan

Takashi Matsuyama

Affiliation Department of Periodontology, Kagoshima University Graduate School of Medical and Dental Sciences, Sakuragaoka, Kagoshima, Japan

Takashi Nishioka

Affiliation Division of Oral Diagnosis, Tohoku University Graduate School of Dentistry, Sendai, Miyagi, Japan

Makoto Hagiwara

Affiliation Department of Oral Disease Research, National Center for Geriatrics and Gerontology, Obu, Aichi, Japan

Yusuke Kiyoura

Affiliation Department of Oral Medical Science, Ohu University School of Dentistry, Koriyama, Fukushima, Japan

Hidetoshi Shimauchi

Affiliation Division of Periodontology and Endodontology, Tohoku University Graduate School of Dentistry, Sendai, Miyagi, Japan

Kenji Matsushita

Affiliation Department of Oral Disease Research, National Center for Geriatrics and Gerontology, Obu, Aichi, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HT KM. Performed the experiments: HT TM TN. Analyzed the data: HT TM TN KM. Contributed reagents/materials/analysis tools: HT TM TN MH YK HS KM. Wrote the paper: HT KM.