About the Authors

Skye Storrie

Affiliation Moredun Research Institute, Pentlands Science Park, Edinburgh, Midlothian, EH26 0PZ, United Kingdom

David Longbottom

Affiliation Moredun Research Institute, Pentlands Science Park, Edinburgh, Midlothian, EH26 0PZ, United Kingdom

Peter G. Barlow

Affiliation School of Life, Sport and Social Sciences, Edinburgh Napier University, Sighthill Campus, Edinburgh, EH11 4BN, United Kingdom

Nick Wheelhouse

n.wheelhouse@napier.ac.uk

Current address: School of Life, Sport and Social Sciences, Edinburgh Napier University, Sighthill Campus, Edinburgh, EH11 4BN, United Kingdom

Affiliation Moredun Research Institute, Pentlands Science Park, Edinburgh, Midlothian, EH26 0PZ, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SS DL PGB NW. Performed the experiments: SS NW. Analyzed the data: SS NW. Contributed reagents/materials/analysis tools: PGB DL. Wrote the paper: SS PGB DL NW.