About the Authors

Kejuan Li

Affiliations Graduate School of Life and Environmental Sciences, University of Tsukuba, 1-1-1 Tennodai, Tsukuba 305–8572, Japan, Drug Discovery and Assets Innovation Laboratory, DAILAB, National Institute of Advanced Industrial Science & Technology (AIST), Central 5–41, 1-1-1 Higashi, Tsukuba 305–8565, Japan, Institute of Laboratory Animal Sciences, Chinese Academy of Medical Sciences, Beijing 100021, China

Yue Yu

Affiliations Graduate School of Life and Environmental Sciences, University of Tsukuba, 1-1-1 Tennodai, Tsukuba 305–8572, Japan, Drug Discovery and Assets Innovation Laboratory, DAILAB, National Institute of Advanced Industrial Science & Technology (AIST), Central 5–41, 1-1-1 Higashi, Tsukuba 305–8565, Japan

Shuang Sun

Affiliation Graduate School of Life and Environmental Sciences, University of Tsukuba, 1-1-1 Tennodai, Tsukuba 305–8572, Japan

Ye Liu

Affiliation Graduate School of Life and Environmental Sciences, University of Tsukuba, 1-1-1 Tennodai, Tsukuba 305–8572, Japan

Sukant Garg

Affiliations Drug Discovery and Assets Innovation Laboratory, DAILAB, National Institute of Advanced Industrial Science & Technology (AIST), Central 5–41, 1-1-1 Higashi, Tsukuba 305–8565, Japan, School of Integrative and Global Majors, University of Tsukuba, 1-1-1 Tennodai, Tsukuba 305–8577, Japan

Sunil C. Kaul

Affiliation Drug Discovery and Assets Innovation Laboratory, DAILAB, National Institute of Advanced Industrial Science & Technology (AIST), Central 5–41, 1-1-1 Higashi, Tsukuba 305–8565, Japan

Zhongfang Lei

Affiliation Graduate School of Life and Environmental Sciences, University of Tsukuba, 1-1-1 Tennodai, Tsukuba 305–8572, Japan

Ran Gao

Affiliation Institute of Laboratory Animal Sciences, Chinese Academy of Medical Sciences, Beijing 100021, China

Renu Wadhwa

renu-wadhwa@aist.go.jp (RW); zhang.zhenya.fu@u.tsukuba.ac.jp (ZZ)

Affiliations Drug Discovery and Assets Innovation Laboratory, DAILAB, National Institute of Advanced Industrial Science & Technology (AIST), Central 5–41, 1-1-1 Higashi, Tsukuba 305–8565, Japan, School of Integrative and Global Majors, University of Tsukuba, 1-1-1 Tennodai, Tsukuba 305–8577, Japan

Zhenya Zhang

renu-wadhwa@aist.go.jp (RW); zhang.zhenya.fu@u.tsukuba.ac.jp (ZZ)

Affiliation Graduate School of Life and Environmental Sciences, University of Tsukuba, 1-1-1 Tennodai, Tsukuba 305–8572, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KL SCK RG RW ZZ. Performed the experiments: KL YY SS SG YL. Analyzed the data: KL YY SG SCK RG RW ZZ. Contributed reagents/materials/analysis tools: SCK RG RW ZZ. Wrote the paper: KL SCK RW ZL ZZ.