About the Authors

Cuihua Gu

Affiliations School of Landscape and Architecture, Zhejiang Agriculture and Forestry University, Hangzhou 311300, P.R. China, Department of Biology, Colorado State University, Fort Collins, Colorado, 80523, United States of America

Luke R. Tembrock

Affiliation Department of Biology, Colorado State University, Fort Collins, Colorado, 80523, United States of America

Nels G. Johnson

Affiliation National Institute for Mathematical and Biological Synthesis, University of Tennessee, Knoxville, 37996, Tennessee, United States of America

Mark P. Simmons

Affiliation Department of Biology, Colorado State University, Fort Collins, Colorado, 80523, United States of America

Zhiqiang Wu

wu.zhiqiang.1020@gmail.com

Affiliation Department of Biology, Colorado State University, Fort Collins, Colorado, 80523, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZW CG. Performed the experiments: ZW CG. Analyzed the data: ZW CG LRT. Contributed reagents/materials/analysis tools: ZW CG LRT. Wrote the paper: ZW CG LRT NGJ MPS.