About the Authors

Alan Kasprak

akasprak@usgs.gov (AK); nate@natehough-snee.org (NH-S)

Current address: U.S. Geological Survey, Grand Canyon Monitoring and Research Center, Flagstaff, AZ, 86001, United States of America

‡ AK and NH-S are joint lead authors on this work. TB, NB, GB, RC, KF, HI, MJ, GO, DR and JW are presented alphabetically on this work.

Affiliation Department of Watershed Sciences, Utah State University, Logan, UT 84322–5210, United States of America

Nate Hough-Snee

akasprak@usgs.gov (AK); nate@natehough-snee.org (NH-S)

‡ AK and NH-S are joint lead authors on this work. TB, NB, GB, RC, KF, HI, MJ, GO, DR and JW are presented alphabetically on this work.

Affiliations Department of Watershed Sciences, Utah State University, Logan, UT 84322–5210, United States of America, Ecology Center, Utah State University, Logan, UT, 84322–5210, United States of America

Tim Beechie

Contributed equally to this work with: Tim Beechie, Nicolaas Bouwes, Gary Brierley, Reid Camp, Kirstie Fryirs, Hiroo Imaki, Martha Jensen, Gary O’Brien, David Rosgen, Joseph Wheaton

‡ AK and NH-S are joint lead authors on this work. TB, NB, GB, RC, KF, HI, MJ, GO, DR and JW are presented alphabetically on this work.

Affiliation Fish Ecology Division, Northwest Fisheries Science Center, National Marine Fisheries Service, National Oceanic and Atmospheric Administration, Seattle, WA 98112, United States of America

Nicolaas Bouwes

Contributed equally to this work with: Tim Beechie, Nicolaas Bouwes, Gary Brierley, Reid Camp, Kirstie Fryirs, Hiroo Imaki, Martha Jensen, Gary O’Brien, David Rosgen, Joseph Wheaton

‡ AK and NH-S are joint lead authors on this work. TB, NB, GB, RC, KF, HI, MJ, GO, DR and JW are presented alphabetically on this work.

Affiliation Eco Logical Research, Providence, UT, United States of America

Gary Brierley

Contributed equally to this work with: Tim Beechie, Nicolaas Bouwes, Gary Brierley, Reid Camp, Kirstie Fryirs, Hiroo Imaki, Martha Jensen, Gary O’Brien, David Rosgen, Joseph Wheaton

‡ AK and NH-S are joint lead authors on this work. TB, NB, GB, RC, KF, HI, MJ, GO, DR and JW are presented alphabetically on this work.

Affiliation School of Environment, University of Auckland, Auckland, New Zealand

Reid Camp

Contributed equally to this work with: Tim Beechie, Nicolaas Bouwes, Gary Brierley, Reid Camp, Kirstie Fryirs, Hiroo Imaki, Martha Jensen, Gary O’Brien, David Rosgen, Joseph Wheaton

‡ AK and NH-S are joint lead authors on this work. TB, NB, GB, RC, KF, HI, MJ, GO, DR and JW are presented alphabetically on this work.

Affiliations Department of Watershed Sciences, Utah State University, Logan, UT 84322–5210, United States of America, Eco Logical Research, Providence, UT, United States of America

Kirstie Fryirs

Contributed equally to this work with: Tim Beechie, Nicolaas Bouwes, Gary Brierley, Reid Camp, Kirstie Fryirs, Hiroo Imaki, Martha Jensen, Gary O’Brien, David Rosgen, Joseph Wheaton

‡ AK and NH-S are joint lead authors on this work. TB, NB, GB, RC, KF, HI, MJ, GO, DR and JW are presented alphabetically on this work.

Affiliation Department of Environmental Sciences, Macquarie University, Sydney, Australia

Hiroo Imaki

Contributed equally to this work with: Tim Beechie, Nicolaas Bouwes, Gary Brierley, Reid Camp, Kirstie Fryirs, Hiroo Imaki, Martha Jensen, Gary O’Brien, David Rosgen, Joseph Wheaton

‡ AK and NH-S are joint lead authors on this work. TB, NB, GB, RC, KF, HI, MJ, GO, DR and JW are presented alphabetically on this work.

Affiliation Pacific Spatial Solutions, Reston, VA, United States of America

Martha Jensen

Contributed equally to this work with: Tim Beechie, Nicolaas Bouwes, Gary Brierley, Reid Camp, Kirstie Fryirs, Hiroo Imaki, Martha Jensen, Gary O’Brien, David Rosgen, Joseph Wheaton

‡ AK and NH-S are joint lead authors on this work. TB, NB, GB, RC, KF, HI, MJ, GO, DR and JW are presented alphabetically on this work.

Affiliations Department of Watershed Sciences, Utah State University, Logan, UT 84322–5210, United States of America, Ecology Center, Utah State University, Logan, UT, 84322–5210, United States of America

Gary O’Brien

Contributed equally to this work with: Tim Beechie, Nicolaas Bouwes, Gary Brierley, Reid Camp, Kirstie Fryirs, Hiroo Imaki, Martha Jensen, Gary O’Brien, David Rosgen, Joseph Wheaton

‡ AK and NH-S are joint lead authors on this work. TB, NB, GB, RC, KF, HI, MJ, GO, DR and JW are presented alphabetically on this work.

Affiliation Department of Watershed Sciences, Utah State University, Logan, UT 84322–5210, United States of America

David Rosgen

Contributed equally to this work with: Tim Beechie, Nicolaas Bouwes, Gary Brierley, Reid Camp, Kirstie Fryirs, Hiroo Imaki, Martha Jensen, Gary O’Brien, David Rosgen, Joseph Wheaton

‡ AK and NH-S are joint lead authors on this work. TB, NB, GB, RC, KF, HI, MJ, GO, DR and JW are presented alphabetically on this work.

Affiliation Wildland Hydrology, Fort Collins, CO, 80524, United States of America

Joseph Wheaton

Contributed equally to this work with: Tim Beechie, Nicolaas Bouwes, Gary Brierley, Reid Camp, Kirstie Fryirs, Hiroo Imaki, Martha Jensen, Gary O’Brien, David Rosgen, Joseph Wheaton

‡ AK and NH-S are joint lead authors on this work. TB, NB, GB, RC, KF, HI, MJ, GO, DR and JW are presented alphabetically on this work.

Affiliations Department of Watershed Sciences, Utah State University, Logan, UT 84322–5210, United States of America, Ecology Center, Utah State University, Logan, UT, 84322–5210, United States of America

Competing Interests

Nicolaas Bowes is the Owner of Eco Logical Research; Hiroo Imaki is the Owner of Pacific Spatial Solutions; David L. Rosgen is the Owner of Wildland Hydrology. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: AK NH-S RC. Performed the experiments: AK NH-S RC. Analyzed the data: AK NH-S RC HI GB TB JW NB GO KF. Contributed reagents/materials/analysis tools: TB HI DR GB KF NH-S GO JW. Wrote the paper: AK NH-S TB NB GB RC KF HI MJ GO DR JW.