About the Authors

Lindsay E. Laurie

Affiliation Research Center for Medical and Dental Sciences, Tokyo Medical and Dental University (TMDU), Bunkyo-ku, Tokyo, Japan

Hiroki Kokubo

Affiliations Division of Mammalian Development, National Institute of Genetics, Mishima, Shizuoka, Japan, Department of Genetics, The Graduate University for Advanced Studies, Mishima, Shizuoka, Japan, Department of Cardiovascular Physiology and Medicine, Graduate School of Biomedical and Health Sciences, Hiroshima University, Minamiku, Hiroshima, Japan

Masataka Nakamura

Affiliation Research Center for Medical and Dental Sciences, Tokyo Medical and Dental University (TMDU), Bunkyo-ku, Tokyo, Japan

Yumiko Saga

Affiliations Division of Mammalian Development, National Institute of Genetics, Mishima, Shizuoka, Japan, Department of Genetics, The Graduate University for Advanced Studies, Mishima, Shizuoka, Japan

Noriko Funato

noriko-funato@umin.ac.jp

Affiliation Research Center for Medical and Dental Sciences, Tokyo Medical and Dental University (TMDU), Bunkyo-ku, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NF. Performed the experiments: LEL HK. Analyzed the data: YS MN. Contributed reagents/materials/analysis tools: YS MN NF. Wrote the paper: LEL NF.