About the Authors

Tatsuya Tada

Affiliation Department of Infectious Diseases, Research Institute, National Center for Global Health and Medicine, 1-21-1 Toyama, Shinjuku, Tokyo, Japan

Tohru Miyoshi-Akiyama

Affiliations Department of Infectious Diseases, Research Institute, National Center for Global Health and Medicine, 1-21-1 Toyama, Shinjuku, Tokyo, Japan, Pathogenic Microbe Laboratory, Research Institute, National Center for Global Health and Medicine, 1-21-1 Toyama, Shinjuku, Tokyo, Japan

Kayo Shimada

Affiliation Department of Infectious Diseases, Research Institute, National Center for Global Health and Medicine, 1-21-1 Toyama, Shinjuku, Tokyo, Japan

Akino Shiroma

Affiliation Research and Development Division, Okinawa Institute of Advanced Sciences, Uruma, Okinawa, Japan

Kazuma Nakano

Affiliation Research and Development Division, Okinawa Institute of Advanced Sciences, Uruma, Okinawa, Japan

Kuniko Teruya

Affiliation Research and Development Division, Okinawa Institute of Advanced Sciences, Uruma, Okinawa, Japan

Kazuhito Satou

Affiliation Research and Development Division, Okinawa Institute of Advanced Sciences, Uruma, Okinawa, Japan

Takashi Hirano

Affiliation Research and Development Division, Okinawa Institute of Advanced Sciences, Uruma, Okinawa, Japan

Masahiro Shimojima

Affiliation Department of Accuracy Management, BML Inc., Kawagoe, Saitama, Japan

Teruo Kirikae

tkirikae@ri.ncgm.go.jp

Affiliation Department of Infectious Diseases, Research Institute, National Center for Global Health and Medicine, 1-21-1 Toyama, Shinjuku, Tokyo, Japan

Competing Interests

M. Shimojima is employed by a commercial company (BML Inc.). This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: TT TK. Performed the experiments: TT TM K. Shimada AS KN KT K. Satou TH. Analyzed the data: TT K. Shimada. Contributed reagents/materials/analysis tools: MS. Wrote the paper: TT TK.