About the Authors

Francesca Mancini

Current address: GlaxoSmithKline Vaccines S.r.l., Siena, Italy

Affiliations Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy, Department of Biomedical Sciences, University of Padua, Padua, Italy

Elisabetta Monaci

Current address: GlaxoSmithKline Vaccines S.r.l., Siena, Italy

Affiliation Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy

Giuseppe Lofano

Current address: Ragon Institute of MGH, MIT and Harvard, Cambridge, Massachusetts, United States of America

Affiliations Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy, Department of Biology and Biotechnologies “Charles Darwin”, University of Rome “La Sapienza”, Rome, Italy

Antonina Torre

Affiliation Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy

Marta Bacconi

Current address: GlaxoSmithKline Vaccines S.r.l., Siena, Italy

Affiliations Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy, Department of Biotechnology, Chemistry and Pharmacy, University of Siena, Siena, Italy

Simona Tavarini

Current address: GlaxoSmithKline Vaccines S.r.l., Siena, Italy

Affiliation Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy

Chiara Sammicheli

Current address: GlaxoSmithKline Vaccines S.r.l., Siena, Italy

Affiliation Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy

Letizia Arcidiacono

Current address: GlaxoSmithKline Vaccines S.r.l., Siena, Italy

Affiliation Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy

Bruno Galletti

Current address: GlaxoSmithKline Vaccines S.r.l., Siena, Italy

Affiliation Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy

Donatello Laera

Current address: GlaxoSmithKline Vaccines S.r.l., Siena, Italy

Affiliation Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy

Michele Pallaoro

Current address: GlaxoSmithKline Vaccines S.r.l., Siena, Italy

Affiliation Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy

Giovanna Tuscano

Current address: GlaxoSmithKline Vaccines S.r.l., Siena, Italy

Affiliation Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy

Maria Rita Fontana

Current address: GlaxoSmithKline Vaccines S.r.l., Siena, Italy

Affiliation Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy

Giuliano Bensi

Current address: GlaxoSmithKline Vaccines S.r.l., Siena, Italy

Affiliation Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy

Guido Grandi

Current address: Centre for Integrative Biology, University of Trento, Trento, Italy

Affiliation Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy

Silvia Rossi-Paccani

Current address: GlaxoSmithKline Vaccines S.r.l., Siena, Italy

Affiliation Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy

Sandra Nuti

Current address: GlaxoSmithKline Vaccines S.r.l., Siena, Italy

Affiliation Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy

Rino Rappuoli

Current address: GlaxoSmithKline Vaccines S.r.l., Siena, Italy

Affiliation Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy

Ennio De Gregorio

Current address: GlaxoSmithKline Vaccines S.r.l., Siena, Italy

Affiliation Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy

Fabio Bagnoli

Current address: GlaxoSmithKline Vaccines S.r.l., Siena, Italy

Affiliation Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy

Elisabetta Soldaini

elisabetta.x.soldaini@gsk.com

Current address: GlaxoSmithKline Vaccines S.r.l., Siena, Italy

Affiliation Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy

Sylvie Bertholet

Current address: GlaxoSmithKline Vaccines S.r.l., Siena, Italy

Affiliation Novartis Vaccines and Diagnostics, S.r.l., Research Center, Siena, Italy

Competing Interests

The authors have read the journal's policy and have the following competing interests: Salaries for all authors were provided by Novartis Vaccines and Diagnostics S.r.l. during completion of the experimental studies contained herein. This does not alter the authors’ adherence to all PLoS One policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: FM EM MRF GB GG RR EDG FB ES SB. Performed the experiments: FM EM GL AT MB LA DL GT SR-P ES. Analyzed the data: FM EM AT GT MRF BG MP SR-P GB SN ES SB. Contributed reagents/materials/analysis tools: ST CS. Wrote the paper: FM FB ES SB.