About the Authors

Bryon A. Nicholson

Affiliation Department of Veterinary Microbiology and Preventive Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Aaron C. West

Affiliation Department of Chemistry, College of Liberal Arts and Sciences, Iowa State University, Ames, Iowa, United States of America

Paul Mangiamele

Affiliation Department of Veterinary Microbiology and Preventive Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Nicolle Barbieri

Affiliation Department of Veterinary Microbiology and Preventive Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Yvonne Wannemuehler

Affiliation Department of Veterinary Microbiology and Preventive Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Lisa K. Nolan

Affiliation Department of Veterinary Microbiology and Preventive Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Catherine M. Logue

Affiliation Department of Veterinary Microbiology and Preventive Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Ganwu Li

liganwu@iastate.edu

Affiliations Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, 1802 University Blvd, Iowa State University, Ames, Iowa, 50011, United States of America, Department of Veterinary Preventive Medicine, College of Veterinary Medicine, Nanjing Agricultural University, Nanjing, Jiangsu, 210095, P. R. China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LKN CML GL YW NB BAN. Performed the experiments: BAN YW. Analyzed the data: BAN ACW. Contributed reagents/materials/analysis tools: ACW PM BAN. Wrote the paper: BAN CML LKN GL.