About the Authors

James P. Charles

Affiliations Neuromuscular Diseases Group, Comparative Biomedical Sciences, Royal Veterinary College, 4 Royal College Street, London, NW1 0TU, United Kingdom, Structure and Motion Lab, Comparative Biomedical Sciences, Royal Veterinary College, Hawkshead Lane, Hatfield, Hertfordshire, AL9 7TA, United Kingdom

Ornella Cappellari

Affiliation Neuromuscular Diseases Group, Comparative Biomedical Sciences, Royal Veterinary College, 4 Royal College Street, London, NW1 0TU, United Kingdom

Andrew J. Spence

Affiliations Structure and Motion Lab, Comparative Biomedical Sciences, Royal Veterinary College, Hawkshead Lane, Hatfield, Hertfordshire, AL9 7TA, United Kingdom, Department of Bioengineering, College of Engineering, Temple University, 1947 N. 12th Street, Philadelphia, PA 19122, United States of America

John R. Hutchinson

Affiliation Structure and Motion Lab, Comparative Biomedical Sciences, Royal Veterinary College, Hawkshead Lane, Hatfield, Hertfordshire, AL9 7TA, United Kingdom

Dominic J. Wells

dwells@rvc.ac.uk

Affiliation Neuromuscular Diseases Group, Comparative Biomedical Sciences, Royal Veterinary College, 4 Royal College Street, London, NW1 0TU, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JRH AJS DJW JPC. Performed the experiments: JPC OC. Analyzed the data: JPC. Contributed reagents/materials/analysis tools: JPC OC DJW. Wrote the paper: JPC OC AJS JRH DJW.